In the 22nd century, scientists from an energy-depleted Earth research new fuel sources in the far corners of the solar system, where they discover an ancient alien race from Jupiter as well as the emergence of an apocalyptically dangerous black hole. Koji Hashimoto's 1983 sci-fi adventure was conceived as a Japanese competitor to the upcoming sequel to Stanley Kubrick's classic 2001, titled 2010, and often mirrors the sequel's plot.
Chief Kuroki tells the team that they will soon be meeting Beet Buster and his new Buddy Roid, but before any introduction can be made, the Soujikiloid attacks. Soon after Hiromu learns that the two new Go-Busters originated from Hyper Space, he challenges them to a duel, suspicious of their motives. This is a director's cut version of episodes 15 & 16 of the Tokumei Sentai Go-Busters TV series.
Japan is thrown into a panic after several ships are sunk near Odo Island. An expedition to the island led by Dr. Kyohei Yamane soon discover something far more devastating than imagined in the form of a 50 meter tall monster whom the natives call Gojira. Now the monster begins a rampage that threatens to destroy not only Japan, but the rest of the world as well.
Two fishing scout pilots make a horrifying discovery when they encounter a second Godzilla alongside a new monster named Anguirus. Without the weapon that killed the original, authorities attempt to lure Godzilla away from the mainland. But Anguirus soon arrives and the two monsters make their way towards Osaka as Japan braces for tragedy.
The communication system of the X-Force base in Kuala Lumpur, Malaysia is hacked by a mysterious saboteur. Bai Lu is assigned to Kuala Lumpur to carry out a reconnaissance mission. The Sky Key, which can open a space-time portal and is stored in the Malaysian base, is stolen. At the same time, Bai Lu suddenly loses contact and is suspected of stealing the Sky Key. She is wanted worldwide. Hong Huan and Huang Li are extremely worried and act rashly without the organization's permission, which unexpectedly causes an uproar.
The advertising director of Pacific Pharmaceuticals, frustrated with the low ratings of their sponsored TV program, seeks a more sensationalist approach. He orders his staff to Faro Island to capture King Kong for exploitation. As Godzilla re-emerges, a media frenzy generates with Pacific looking to capitalize off of the ultimate battle.
Journalists Ichiro Sakai and Junko cover the wreckage of a typhoon when an enormous egg is found and claimed by greedy entrepreneurs. Mothra's fairies arrive and are aided by the journalists in a plea for its return. As their requests are denied, Godzilla arises near Nagoya and the people of Infant Island must decide if they are willing to answer Japan's own pleas for help.
A special retrospective of the two franchises, with segments making comparisons between the different characters, their weapons, powers and so forth. The final segment of the special is the short film "Super Battle: Ultraman vs Kamen Rider" The main characters, Ultraman and Kamen Rider 1 appear on the scene separately, each fighting an original kaijin and kaiju. When the two monsters are nearly defeated, they merge into a more powerful kaiju. Ultraman experiences trouble and calls for Kamen Rider 1's help. Kamen Rider 1 is then able to grow to enormous size to fight alongside Ultraman.
In the spring of 1975, peace returned to the universe once more. But then members of the Ultra Brothers started being assasinated by someone, one by one. The perpetrator was Jackal the Demonic Space Overlord. Jackal was an alien defeated by Ultraman King and banished to spend an eternity in a black hole. Having returned, Jackal sent his armies to Earth to uncover any Ultra Warriors that may be hiding in humanoid form there. Jackal's minions resorted to threatening to kill human beings held as hostages in order to force Ultra Warriors to reveal themselves. Just when it seemed all hope was lost, a mysterious warrior wearing a suit of armor that obscured his identity appeared from the heavens...
Zero finds himself trapped within a universal gap and faced against an impostor Ace. Following the battle, Darklops Zero was restrained once more by Herodia, remembering a figure named "Kaiser Belial" before deactivating. Rei was restrained after his failed attempt to intrude the alien base but was rescued by none other than his alternate self. Unlike the prime Rei, alternate Reimon was never able to control his powers, forced into the state of Reionic Burst until the original Rei punched Reimon, thus deactivating its Reionic Burst mode. Rei revealed that he lost his Rayblood instinct after Hyuga threw Seven's Eye Slugger, creating a scar on his chest which symbolizes the former's friendship with ZAP SPACY members. While searching for Rei, Hyuga instead finds his Neo Battle Nizer and an entrance to the alien base made by Litra. Although the fake Ace was destroyed, Zero try to find an escaped but was halted by Seven and Ultraman impostors.
After a fishing boat is attacked, the sole surviving crew member realizes it is none other than a resurrected Godzilla. However, efforts to bring the story to light are suppressed by the Japanese government amid growing political tensions between the United States and the Soviet Union, who are both willing to bomb Japan to stop the monster.